Jakub Prokop

   Ph. D. Charles University, Praha

   Phone: ++420/221951837; Fax: ++420/221951841
   E-mail: jprokop@natur.cuni.cz


Research interests
Palaeoentomology; comparative and functional morphology; taxonomy and
phylogeny of Paleozoic insects, mainly Palaeodictyopterida (Palaeodictyoptera,
Megasecoptera, Diaphanopterodea, Permothemistida), Panephemeroptera,
Odonatoptera; wing venation; insects in amber inclusions and sedimentary rocks;
plant-insect interactions in the fossil record.

The functional morphology and evolution of keystone body parts in early
diverging lineages of winged insects (2018-2020)

Lectures and courses
Basic entomology (lectures); Invertebrate zoology (lectures); Entomology
(lectures, practices, field trip),  Advanced entomology (lectures, practices),
System and phylogeny of invertebrates (field trip)

Postdoctoral fellow
Martina Pecharová PhD.

Graduate students
Jan Batelka MSc.
Tomáš Dvořák MSc.
Dominika Ružičková MSc.
Michal Tkoč MSc.

Undergraduate students
Kateřina Rosová
Šárka Škorpíková

Publications (via Google Schoolar, Researchgate,

Batelka J., Prokop J., Pohl H., Bai M., Zhang W. & Beutel R.G. in press. Highly specialized Cretaceous beetle parasitoids (Ripiphoridae) identified with optimized visualization of microstructures. Systematic entomology doi: 10.1111/syen.12331


Batelka J., Engel M.S. & Prokop J. 2018. A remarkable diversity of parasitoid beetles in Cretaceous amber, with a summary of the Mesozoic record of Tenebrionoidea. Cretaceous Research 90: 296–310.

Jacquelin L., Desutter-Grandcolas L., Chintauan-Marquier I., Boistel R., Zheng D., Prokop J., & Nel A. 2018. New insights on basivenal sclerites using 3D tools and homology of wing veins in Odonatoptera (Insecta). Scientific Reports 8: 238, doi: 10.1038/s41598-017-18615-0

Nel A, Prokop J., Pecharová M., Engel M.S. & Garrouste R. 2018. Palaeozoic giant dragonflies were hawker predators. Scientific Reports 8: 12141. doi 10.1038/s41598-018-30629-w

Nidergas V., Hadrava J., Garrouste R., Prokop J., Schubnel T. & Nel A. 2018. The first pipizine hoverfly genus and species from the Oligocene of France (Diptera: Syrphidae). Acta Palaeontologica Polonica 63 (3): 539–554.

Pohl H., Batelka J., Prokop J., Müller P., Yavorskaya M.I. & Beutel R.G. 2018. A needle in a haystack: Mesozoic origin of parasitism in Strepsiptera revealed by first definite Cretaceous primary larva (Insecta) Insecta). PeerJ 6:e5943 doi: 10.7717/peerj.5943

Pecharová M. & Prokop J. 2018. The morphology of mouthparts, wings and genitalia of Paleozoic insect families Protohymenidae and Scytohymenidae reveals new details and supposed function. Arthropod Structure & Development 47 (1): 117–129.

Prokop J., Pecharová M., Jarzembowski E.A. & Ross A.J. 2018. New palaeodictyopterans from the Late Carboniferous of the UK (Insecta: Palaeodictyopterida). Earth and Environmental Science Transactions of the Royal Society of Edinburgh 107 (2–3): 99–107.

Prokop J., Pecharová M., Nel A. & Hörnschemeyer T. 2018. The wing base of the palaeodictyopteran genus Dunbaria Tillyard: Where are we now? Arthropod Structure & Development 47 (4): 339-351.


Prokop J., Pecharová M., Garrouste R., Beattie R., Chintauan-Marquier I.C. & Nel A. 2017. Redefining the extinct orders Miomoptera and Hypoperlida as stem acercarian insects. BMC Evolutionary Biology (2017) 17:205 doi: 10.1186/s12862-017-1039-3

Prokop J. & Kukalová-Peck J. 2017. New insects from the earliest Permian of Carrizo Arroyo (New Mexico, U. S. A.) bridging the gap between Carboniferous and Permian entomofaunas. Insect Systematics & Evolution 48: 493–511.

Prokop J., Dehon M., Michez D. & Engel M.S. 2017. An Early Miocene bumble bee from northern Bohemia (Hymenoptera: Apidae). ZooKeys 710: 43–63.

Prokop J., Pecharová M., Nel A., Grey M. & Hörnschemeyer T. 2017. A remarkable insect from the Pennsylvanian of the Joggins Formation in Nova Scotia, Canada: insights into unusual venation of Brodiidae and nymphs of Megasecoptera. Journal of Systematic Palaeontology 15(12): 1051–1065. doi: 10.1080/14772019.2017.1283364

Prokop J., Pecharová M., Nel A., Hörnschemeyer T., Krzemińska E., Krzemiński W. and Engel M. S. 2017. Paleozoic nymphal wing pads support dual model of insect wing origins. Current Biology 27(2): 263–269. 


Prokop J., Pecharová M. & Ren D. 2016. Hidden surface microstructures on Carboniferous insect Brodioptera sinensis (Megasecoptera) enlighten functional morphology and sensorial perception. Scientific Reports 6, 28316; doi: 10.1038/srep28316

Prokop J., Pecharová M. & Nel A. 2016. New Cenozoic dragonflies from the Most Basin and Středohoří Complex volcanic area (Czech Republic, Germany). Journal of Natural History 50 doi: 10.1080/00222933.2016.1193648

Prokop J, Garrouste R. & Nel A. 2016. A new permopsocidan genus and species from the Late Permian of Australia (Insecta: Acercaria: Psocidiidae). Zootaxa 4147(5):589. doi: 10.11646/zootaxa.4147.5.7

Prokop J., Nel A., Engel M.S., Pecharová M. & Hörnschemeyer T. 2016. New Carboniferous fossils of Spilapteridae enlighten postembryonic wing development in Palaeodictyoptera. Systematic entomology 41: 178–190.

Tkoč M., Nel A. & Prokop J 2016. Discovery of a new species of the Cretaceous genus Microphorites Hennig, 1971 (Diptera: Dolichopodidae s. lat.) in Paleogene amber from eastern Moravia (Czech Republic). Insect Systematics & Evolution 47 (2): 181–193.

Guan Z., Prokop J., Roques P., Lapeyrie J. & Nel A. 2016. Revision of the enigmatic insect family Anthracoptilidae enlightens the evolution of Palaeozoic stem-dictyopterans. Acta Palaeontologica Polonica 61 (1): 71–87.

Huang D.Y, Bechly G., Nel P., Engel M.S., Prokop J., Azar D., Cai C.Y., van de Kamp T., Staniczek A., Garrouste R., Krogmann L., dos Santos Rolo T., Baumbach T., Ohlhoff R., Shmakov A.S., Bourgoin T. &Nel A. 2016. New fossil insect order Permopsocida elucidates major radiation and evolution of suction feeding in hemimetabolous insects (Hexapoda: Acercaria). Scientific Reports 6, 23004 doi: 10.1038/srep23004


Pecharová, M., Prokop, J. & Ren D. 2015. Early Pennsylvanian aykhalids from Xiaheyan of northern China and their palaeogeographical significance (Insecta: Megasecoptera). Comptes Rendus Palevol 14: 613–624.

Prokop J., Szwedo J., Lapeyrie J., Garrouste R. & Nel A 2015. New middle Permian insects from Salagou Formation of the Lodève Basin in southern France (Insecta: Pterygota). Annales de la Société Entomologique de France 51(1): 14–51.

Fernandes F.R., Prokop J., & Nel A. 2015. Discovery of the genus Crinoedischia in the middle Permian of France (Insecta: Orthoptera). Annales de la Société Entomologique de France 50(3-4): 260–263.

Knor S., Kvaček Z., Wappler T. & Prokop J. 2015 Diversity, taphonomy and palaeoecology of plant–arthropod interactions in the Lower Miocene (Burdigalian) of the Most Basin in north–western Bohemia. Review of Palaeobotany and Palynology 219 (2015): 52–70

Nel A., Garrouste R., & Prokop J 2015. The first African Anthracoptilidae (Insecta: Paoliida) near the Permian–Triassic boundary in Kenya. Zootaxa 3925 (1): 145–150.

Pecharová, M., Ren D. & Prokop, J. 2015. A new palaeodictyopteroid (Megasecoptera: Brodiopteridae) from the Early Pennsylvanian of northern China reveals unique morphological traits and intra-specific variability. Alcheringa 39(2): 236–249.


Coty D., Háva J., Prokop J., Roques P. & Nel A. 2014. New archaeorthopteran insects from the Late Carboniferous of the Nord and Pas-de-Calais basins in northern France (Insecta: Cnemidolestodea, Panorthoptera). Zootaxa 3878(5): 462–470.

Kirejtshuk A, Poschmann M., Prokop J., Garrouste R. & Nel A. 2014. Evolution of the elytral venation and structural adaptations in the oldest Palaeozoic beetles (Insecta: Coleoptera: Tshekardocoleidae). Journal of Systematic Palaeontology 12(5): 575–600.

Nel A. , Prokop J., Grandcolas P., Garrouste R., Lapeyrie J., Legendre F., Anisyutkin L.N. & Kirejtshuk A. 2014. The beetle-like Palaeozoic and Mesozoic roachoids of the so-called “umenocoleoid” lineage (Dictyoptera: Ponopterixidae fam. nov.). Comptes Rendus Palevol 13: 545–554.

Prokop J., Krzemiński W., Krzemińska E. Hörnschemeyer T. Ilger J.M., Brauckmann C., Grandcolas P. & Nel A. 2014. Late Palaeozoic Paoliida is the sister group of Dictyoptera (Insecta: Neoptera). Journal of Systematic Palaeontology 12(5): 601–622.

Prokop J., Roques P. & Nel A. 2014. New non-holometabolous insects from Pennsylvanian of Avion locality in Pas-de-Calais, France (Insecta: ‘Exopterygota’). Alcheringa 38: 155–169.

Wappler T., Dlussky G.M., Engel M.S, Prokop J. & Knor S. (2014). A new trap-jaw ant species of the genus Odontomachus (Hymenoptera: Formicidae: Ponerinae) from the Early Miocene (Burdigalian) of the Czech Republic. Paläontologische Zeitschrift 88: 495–502.


Engel M.S., Davis S.R. & Prokop J. 2013. Chapter XII: Insect wings: The evolutionary development of Nature's first flyers. Pp. 269–298. In: Minelli A., Boxshall G. & Fusco G. (eds): Arthropod Biology and Evolution - Molecules, Development, Morphology. Springer

Knor S., Skuhravá M., Wappler T. & Prokop J. 2013. Galls and gall makers on plant leaves from the Early Miocene (Bilina mine, Czech Republic): systematic and palaeoecological implications. Review of Palaeobotany and Palynology 188: 38–51.

Li Y., Ren D., Pecharová M., & Prokop J. 2013. A new palaeodictyopterid from the Upper Carboniferous of China enlightens close relationships of insect fauna between Quilianshian (northern China) and Laurussia (Insecta: Palaeodictyoptera: Spilapteridae). Alcheringa 37(4): 487–495.

Nel, A., Roques, P., Nel, P., Prokin, A.A., Bourgoin, T., Prokop, J., Szwedo, J., Azar, D., Desutter-Grandcolas, L., Wappler, T., Garrouste, R., Coty, D., Huang, D., Engel, M.S. & Kirejtshuk, A.G. 2013. The earliest known holometabolous insects. Nature 503: 257–261.

Prokop J., Tippeltová Z., Roques P. & Nel A. 2013. A new genus and species of Breyeriidae and wings of immature stages from the Upper Carboniferous, Nord-Pas-de-Calais, France (Insecta: Palaeodictyoptera). Insect Systematics & Evolution 44: 117–128.


Nel A., Ilger J.M., Brauckmann C. & Prokop J. 2012. Revision of Bechala sommeri Ilger and Brauckmann, 2012 from the early Late Carboniferous of Germany (Insecta: Odonatoptera: Bechalidae). Insect Systematics & Evolution 43: 161169.

Fikáček M., Prokin A., Angus R.B., Ponomarenko A., Yue Y., Ren D. & Prokop J. 2012. Phylogeny and the fossil record of the Helophoridae reveal Jurassic origin of modern hydrophiloid lineages (Coleoptera: Polyphaga). Systematic Entomology 37: 420447.

Fikáček M., Prokin A., Angus R.B., Ponomarenko A., Yue Y., Ren D. & Prokop J. 2012. Revision of Mesozoic fossils of the helophorid lineage of the superfamily Hydrophiloidea (Coleoptera: Polyphaga). Acta Entomologica Musei Nationalis Pragae 52(1): 89–127.

Prokop J., Weiß K.D., Dechambre R.-D. & Nel A.2012. Early Permian insects from Saar–Nahe Basin of Odernheim town site, Rheinland-Pfalz in Germany (Insecta: Grylloblattida, Blattinopseida). Geodiversitas 34 (2): 271-281.

Nel P., Azar D, Prokop J., Roques P., Hodebert G. & Nel A. 2012. From Carboniferous to Recent: wing venation enlightens evolution of thysanopteran lineage. Journal of Systematic Palaeontology 10(2): 385-399.

Nel A., Prokop J., Nel P., Grandcolas P., Huang D.-Y., Roques P., Guilbert E., Dostál O. & Szwedo J. 2012. Traits and evolution of wing venation pattern in paraneopteran insects. Journal of Morphology 273(5): 480–506.

Prokop J., Krzemiński W., Krzemińska E. & Wojtechowski D. 2012. Paoliida, a putative stem-group of winged insects: Morphology of new taxa from the Upper Carboniferous of Poland. Acta Palaeontologica Polonica 57(1): 161-173.

Knor S., Prokop J., Kvaček Z., Janovský Z. & Wappler T. 2012. Plant-Arthropod associations from the Early Miocene of the Most Basin in North Bohemia – Palaeoecological and palaeoclimatological implications. Palaeogeography, Palaeoclimatology, Palaeoecology 321-322(2012): 102-112.

Nel A., Bechly G., Prokop J., Béthoux O. & Fleck G. 2012. Systematics and evolution of Palaeozoic and Mesozoic damselfly-like Odonatoptera of the ‘protozygopteran’ grade. Journal of Palaeontology 86(1): 81-104.


Krzeminski W. & Prokop J. 2011. Ptychoptera deleta Novák, 1877 from the Early Miocene of the Czech Republic: redescription of the first fossil attributed to Ptychopteridae (Diptera). ZooKeys 130: 299-305.

Prokop J. & Nel A. 2011. New Middle Permian palaeopteran insects from Lodève Basin in southern France (Insecta: Ephemeroptera, Diaphanopterodea, Megasecoptera). ZooKeys 130: 41-55.

Batelka J., Engel M. S., Falin Z.H. & Prokop J. 2011. Aggregative behaviour of ripidiine males “frozen” in the fossil record, with the description of two new species in Dominican Amber (Coleoptera: Ripiphoridae). European Journal of Entomology 108: 275-286.


Azar D., Prokop J., & Nel A. 2010. The first damselfly from Lebanese amber (Odonata, Zygoptera, Lestomorpha). Alavesia 3: 73-79.

Prokop J., Wappler T., Knor S. & Kvaček Z. 2010. Plant-arthropod associations from the Lower Miocene of the Most Basin in northern Bohemia (Czech Republic): A preliminary report. Acta Geologica Sinica 84(4): 903-914.

Fikáček M., Prokop J. & Nel A. 2010. Fossil water scavenger beetles of the subtribe Hydrobiusina (Coleoptera: Hydrophilidae) from the Late Oligocene locality of Aix-en-Provence. Acta Entomologica Musei Nationalis Pragae 50: 445-458.

Fikáček M., Heiko S & Prokop J. 2010. Fossil hydrophilid beetles (Coleoptera: Hydrophilidae) f the Late Oligocene Rott Formation (Germany). Acta Geologica Sinica 84(4): 732-750.

Prokop J., Nel A. & Tenny A. 2010. On the phylogenetic position of the palaeopteran Syntonopteroidea (Insecta: Ephemeroptera), with a new species from the Upper Carboniferous of England. Organisms Diversity & Evolution 10: 331-340.

Nel A., Nel P., Petrulevičius J.F., Perrichot V., Prokop J., & Azar D. 2010. The Wagner Parsimony using Morphological Characters: a new method for palaeosynecological studies. Annales de la Société Entomologique de France 46(1-2): 276-292.

Prokop J., & Nel A. 2010. New griffenfly, Bohemiatupus elegans from the Late Carboniferous of western Bohemia in the Czech Republic (Odonatoptera: Meganisoptera: Meganeuridae). Annales de la Société Entomologique de France 46(1-2): 183-188.


Nel A., Fleck G., Garrouste R., Gand G., Lapeyrie J., Bybee S.M. & Prokop J. 2009. Revision of Permo-Carboniferous griffenflies (Insecta: Odonatoptera: Meganisoptera) based upon new species and redescription of selected poorly known taxa from Eurasia. Palaeontographica Abt. A 289: 89-121.

Nel A. & Prokop J. 2009. Go to the species in paleoentomology: illusion or necessity. Mémoires de la Société Entomologique de France 8 (2009): 49-55.

Prokop J. & Nel A.2009 Systematic position of Triplosoba, hitherto the oldest mayfly from Upper Carboniferous of Commentry in Central France (Insecta: Palaeodictyopterida). Systematic Entomology 34: 610-615. [IF 1.808]

Dostál O. & Prokop J. 2009 New fossil insects from the Lower Permian of Boskovice Basin in southern Moravia (Diaphanopterodea: Martynoviidae). Geobios 42(4): 495-502.

Liu Yu., Ren D. & Prokop J. 2009. Discovery of a new Namurian archaeorthopterid from Ningxia, China (Insecta: Archaeorthoptera). Zootaxa 2032: 63–68.


Háva J., Prokop J.& Herrmann A. 2008. New fossil dermestid beetles (Coleoptera: Dermestidae) from the Baltic amber - III. Acta Societatis Zoologicae Bohemicae 71(3-4): 151–157.

Fikáček, M., Hájek J. & Prokop J. 2008. New records of the water beetles (Coleoptera: Dytiscidae, Hydrophilidae) from the central European oligocene-miocene deposits, with a confirmation of the generic status of  Hydrobiomorpha enspelense Wedmann, 2000. Annales de la Société Entomologique de France 44(2): 187-199.

Nel A., Prokop J. & Ross A. J. 2008. New genus of leaf-mimicking katydids (Orthoptera: Tettigoniidae) from the late Eocene – early Oligocene of France and England. Comptes Rendus Palevol 7(4): 211-216.

Ren, D., Nel A. & Prokop J. 2008. New early griffenfly, Sinomeganeura huangheensis from the Late Carboniferous of northern China (Meganisoptera: Meganeuridae). Insect Systematics & Evolution 38: 223-229.


Prokop J. & Fikáček M. 2007. Early Oligocene insect fauna from Seifhennersdorf (Saxony, Germany). Acta Musei Nationalis Pragae, Series B, Historia Naturalis 63(2-4): 209-217.

Nel A., Roques P., Nel P., Prokop J. & Steyer J.S. 2007. The earliest holometabolous insect from the Carboniferous: a “crucial” innovation with delayed success (Insecta Protomeropina Protomeropidae). Annales de la Société Entomologique de France 43(3): 349-355.

Prokop J. & Nel A. 2007. New fossil record Gerridae (Heteroptera: Gerromorpha) from the Early Oligocene diatomite in the Czech Republic. Acta Societatis Zoologicae Bohemmicae 71 (1): 15-19.

Prokop J. & Nel A.2007. An enigmatic Palaeozoic stem-group: Paoliida, designation of new taxa from the Upper Carboniferous of the Czech Republic (Insecta: Paoliidae, Katerinkidae fam. n.). African Invertebrates 48 (1): 77-86.

Prokop J. & Ren D.2007. New significant fossil insects from the Upper Carboniferous of Ningxia in northern China (Insecta: Palaeodictyoptera, Archaeorthoptera). European Journal of Entomology 104(2): 267-275.

Prokop J., Přikryl T., Dostál O. & Nel A. 2007. Oligaeschna kvaceki sp. nov., a new fossil dragonfly (Odonata: Aeshnidae) from the Middle Oligocene of northern Moravia (West Carpathians). Geologica Carpathica 58(2): 181-184.


Prokop J., Smith R., Jarzembowski E. & Nel A. 2006. New homoiopterids from the Late Carboniferous of England (Insecta: Palaeodictyoptera). C.R. Palevol 5: 867-873.

Háva J. &  Prokop J.2006. Trinodes puetzi sp. nov., a new fossil species described from the Baltic Amber (Coleoptera: Dermestidae). Acta Societatis Zoologicae Bohemicae 69(3-4): 277-279.

Háva J., Prokop J.& Herrmann A. 2006. New fossil dermestid beetles (Coleoptera: Dermestidae) from the Baltic amber. Acta Societatis Zoologicae Bohemicae 69(3-4): 281- 287.

Háva J., Prokop J. & Kadej M. 2006. New fossil dermestid beetles (Coleoptera: Dermestidae) from the Baltic amber – II. Studies and reports of District Museum Prague-East, Taxonomic Series 2(1-2): 1-14.

Nel A. & Prokop J. 2006. New fossil gall midges from the earliest Eocene French amber (Insecta, Diptera, Cecidomyiidae). Geodiversitas 28(1): 37-54.


Prokop J. & Nel A. 2005. New scuttle flies from the Paleogene amber in eastern Moravia of the Czech Republic (Diptera: Phoridae). Studia dipterologica 12(1): 13-22.

Nel A. & Prokop J. 2005. New fossil Scelionidae (Insecta: Hymenoptera) in early Paleogene amber from eastern Moravia (Czech Republic) and northern France. Polskie Pismo Entomologiczne 74: 339-347.

Nel A. Prokop J. De Ploëg G. Millet J. 2005. New Psocoptera (Insecta) from the lowermost Eocene amber of Oise, France. Journal of Systematic Palaeontology 3(2005): 371-391.

Prokop J., Nel A. & Hoch I. 2005: Discovery of the oldest known Pterygota in the Lower Carboniferous of the Upper Silesian Basin in the Czech Republic (Insecta: Archaeorthoptera). Geobios 38(3): 383-387.


Háva J. & Prokop J. 2004. New fossil dermestid-beetles (Coleoptera: Dermestidae) from the Dominican amber of the Greater Antilles, with an appendix listing known fossil species. Acta Societatis Zoologicae Bohemicae 68: 173-182.

Kvaček Z., Böhme M., Dvořák Z., Konzalová M., Mach K., Prokop J. & Rajchl M. 2004: Early Miocene freshwater and swamp ecosystems of the Most Basin (northern Bohemia) with particular reference to the Bílina mine section [Spodnomiocenní sladkovodní a bažinné ekosystémy mostecké pánve (severní Čechy) se zvláštním zřetelem k profilu dolu Bílina]. Journal of the Czech Geological Society 49 (1-2): 1-40.

Nel A. & Prokop J. 2004: New Paleogene Scatopsidae (Diptera: Nematocera) in amber from eastern Moravia and northern France. Acta Societatis Zoologicae Bohemicae, 68: 91-98.

Prokop J. & Nel A. 2004: A new genus and species of Homoiopteridae from the Upper Carboniferous of the Intra-Sudetic Basin, Czech Republic (Insecta: Palaeodictyoptera). European Journal of Entomology 101(4): 583-589.

Prokop J., Nel A., Hájek J. & Bubík M. 2004: First record of a fossil beetle (Coleoptera, Haliplidae) from the base of Paleocene flysch sediments in the Magura Unit (Outer Western Carpathians, Moravia). Geologica Carpathica 55(6): 469-473.


Prokop J.2003: Remarks on palaeoenviromental changes based on reviewed Tertiary insect associations from the Krušné hory piedmont basins and the České středohoří Mts. in northwestern Bohemia (Czech Republic). Acta Zoologica Cracoviensia, 46(suppl. – Fossil Insects): 329-344.

Prokop J. 2003: Jantarová okna do minulosti. Vesmír 82: 682-691.

Prokop J., Fleck G. & Nel A. 2003. New dragonflies from the Lower Miocene (Ottnagian/Karpatian) of the Cypris Shale in western Bohemia (Odonata: Libellulidae). Neues Jahrbuch für Geologie und Paläontologie, Monatshefte, 2003(9): 561-576.

Prokop J. & Nel A. 2003. New fossil Aculeata from the Oligocene of the České středohoří Mts and the Lower Miocene of the Most Basin in northern Czech Republic (Hymenoptera: Apidae, Vespidae). Acta Musei Nationalis Pragae, Series B, Historia Naturalis, 59(3-4): 91-98 + 1 pl.


Prokop J. 2002. An immature stonefly from Lower Miocene of the Bílina mine in northern Bohemia (Plecoptera: Perlidae). - Acta Societatis Zoologicae Bohemicae, 66. 235-239.

Prokop J. & Nel A. 2002. New Tertiary dragonflies from Lower Oligocene of the České středohoří Mts and Lower Miocene of the Most Basin in the Czech Republic (Odonata: Anisoptera). Acta Societatis Zoologicae Bohemmicae, 66: 141-150.


Prokop J. & Nel A. 2000. New fossil dragonflies from the Lower Miocene of the northern Bohemia (Odonata: Aeshnidae). European Journal of Entomology 97(3): 427-431.

Prokop J. & Nel A. 2000.  First record of the genus Lethocerus Mayr, 1853 from the Lower Miocene of the Most Formation in northern Bohemia, Czech Republic (Heteroptera: Nepomorpha: Belostomatidae). Bulletin de la Société Entomologique de France 105(5): 491-495.

Prokop J. & Boulard M. 2000. Tibicina sakalai n. sp., Cigale fossile du Miocène inférieur de Republique tchèque (Auchenorhyncha, Cicadidae Tibicinae). EPHE, Biologie et Évolution des Insectes 13 (2000): 127-131.


Prokop J. & Bílý S. 1999. Dicerca bilinica sp. n. a new species of Buprestid-beetle (Coleoptera: Buprestidae) from Lower Miocene of the Most Formation in northern part of the Czech Republic. Acta Societatis Zoologicae Bohemicae 63: 311-314.

Prokop J. & Nel A. 1999.  Tertiary termite from Bilina Mine locality in northern Bohemia (Isoptera: Hodotermitidae). Klapalekiana 35: 141-144.

Prokop J. 1999: Nové nálezy hmyzu v neogénu Českého masívu. Živa 1: 32-33.